The TRY Database

The TRY database currently comprises about 65 datasets (about 225 contributed files) from more than 90 contributors. It contains about 2.400.000 trait entries for more than 64.000 plant species and about 1000 different traits, covering a variety of biomes and geographic areas. Each trait entry is accompanied by several auxiliary data. Among these datasets are small contributions as well as several large datasets that are already based on joint initiatives and which cover several thousands of entries. These different datasets are gathered in one consistent format in the TRY database. The database construction is under the technical responsibility of the Group for Organismic Biogeochemistry at the Max Planck Institute for Biogeochemistry in Jena, Germany.

Datasets contributed to TRY: TRY_db_Datasets_2009_07.pdf


Location of sites

  • > 1 million trait entries geographically referenced
  • 8161 sites

Species

The original text strings of contributed species names are imported into the TRY database. A copy is reduced to binomial taxonomy omitting non-valid text elements. Binomial names were compared (based on Levenshtein distance) to species nomenclature based on IPNI and Tropicos. After standardization of species names the TRY database contains trait data for about 64.000 species.

List of species names in TRY: TRY_db_Species_2009_07.xls


Traits

Below please find the number of trait entries, geographically referenced entries, species characterized, maps with the location of sites and charts presenting the range of trait values (for selected traits only).

List of database entries by traits: TRY_db_Entries_2009_07.xls
List of traits by species: Download from FTP server

DataIDTraitMapChartEntriesGeoRefSpecies
      
 All entries 2462562105897464000
1Leaf area31461233115129
2Leaf toughness38426072888
3Leaf angle 41450414194392
4Wood density2730386089129
5Wood anatomy  000
6Bifurcation ratio 363636
7Rooting depth 455280443
8Mycorrhizal type 14935104812453
9Nitrogen-fixation capacity 31592174689869
10Root/shoot ratio729168100
11Leaf absorbance  3630137
12Specific leaf area (SLA)50956287547935
13Leaf lifespan (longevity)182515551017
14Leaf carbon content per dry mass816763682080
15Leaf nitrogen content per dry mass37581285766395
16Leaf phosphorus content per dry mass19323140244324
17Leaf distribution along the axis 1075113404136
18Leaf compoundness 1789961087944
19Typical plant height759047324027
20Maximum plant height2093868979222
21Leaf mesophyll cell size (volume) 94694
22Crown area  662
24Stem diameter at 1.3 m (DBH)20992099839
25Photosynthetic pathway 34752517215690
27Resprouting capacity after disturbance 25583672471
28Bark thickness18318352
29Root metamorphoses 417435581535
30Seed dry mass2733078249194
31Seed length973501998
32Dispersal syndrom 2578751607653
33Pollination syndrome 163675794149
34Tolerance to drought  32332812809
35Tolerance to frost  000
36Tolerance to high temperatures  496502740
37Seed longevity  333
38Germination stimulation  9631648
39Germination stimulation by digestion  111
40Tolerance to windthrow  000
41Surface roughness  31031
42Leaf phenology type 578343326414211
43Woodiness 417402352914819
44Potential decomposition rate of litter 14351099913
45Photosynthesis per leaf dry mass340218861231
46Leaf respiration per dry mass2615973530
47Plant growth form 808942850327522
48Leaf type 1411441894355
49Leaf K content per dry mass416420491417
50Stomatal conductance per leaf area15891182693
52Leaf area (leaflets) 66740657476805
53Leaf thickness 334422141680
54Leaf water content 35533374336
55Leaf density 14631140546
56Leaf mass per area based on fresh weight 154094
57CO2 drawdown 1 371371347
58Intercellular CO2 concentration 1627485508
64Leaf mass per area (LMA) 000
65Leaf nitrogen content per area 13843101363748
66Leaf phosphorus content per area 538446262032
67Leaf K content per area 240240218
68Photosynthesis per leaf area 447622181300
69Leaf respiration per area 18161052533
70Leaf respiration per dry mass at growth temperature 535535246
71Leaf respiration per dry mass at 25 C 1654535270
72Leaf respiration per dry mass measured at 25 C 268268239
73Leaf dry mass 844075112194
74Whole leaf dry mass 1796487161908
75ca_ci_2 498498448
76CO2 drawdown 2 507507457
100Leaf nitrogen/phosphorus (N/P) ratio 1180975282982
101Whole leaf nitrogen content 10061006849
102Whole leaf phosphorus content 444444381
105Plant functional type (Sheffield DGVM 1)  254825481981
106Plant functional type (Sheffield DGVM 2)  254825481981
107Plant functional type (Sheffield DGVM 3)  254825481981
108Plant functional type (Biome-BGC 1)  158115811200
109Plant functional type (Biome-BGC 2)  217021701628
110Plant functional type (LPJ DGVM 1)  254825481981
111Plant functional type (LPJ DGVM 2)  254825481981
115Species phylogenic group  617735082651
177Seed mass 34111446813115
190Maximum plant longevity  1477235097556
208Tolerance to shade  30431002549
209Tolerance to waterlogging  8060801
214Leaf inrolling of lamina  598567524
216Stomata density  367093
217Stomata distribution  141062
219Thorn / spine length  598567524
224Root architecture  306280279
226Leaf phenology 2 (seasonal timing of foliage)  31925672896
228Seed terminal velocity 1217275628
232Stomatal index (stomata / epidermis cells)  107055
233Stomatal area  109057
234Stomatal area per one sided leaf area  56056
245"Litter mass loss decomposability"  604592204
248Litter SLA 400400126
249Litter nitrogen content per dry mass  223223116
250Litter phosphorus content per dry mass  999938
251Litter carbon content per dry mass  214214109
252Litter lignin content per dry mass  255153132
253Litter tannin content per dry mass  11011044
254Litter water soluble carbon content  666
256Litter cellulose content  245143125
257Relative growth rate (RGR)  2412317829
258Leaf dry matter content (LDMC) 1735799252496
268Leaf delta 15nitrogen 11115108201679
270Leaf delta 15nitrogen - soil delta 15nitrogen 36643664565
272Root nitrogen content per dry mass 1263158215
273Root delta 15nitrogen  441148117
276Root density  2920101
277Root diameter  2930101
278Root carbon content per dry mass 37817127
279Leaf soluble components content per dry mass  2930101
280Root soluble components content per dry mass  2930101
281Leaf lignin content per dry mass  761468328
282Root lignin content per dry mass  2930101
283Leaf delta13carbon  579284349
284Root delta13carbon  2930101
285Root cellulose content per dry mass  2930101
286Leaf cellulose content per dry mass  2930101
287Root hemi-cellulose content per dry mass  2930101
288Leaf hemi-cellulose content per dry mass  2930101
289Dispersal syndrom Animal group  196201771
290Dispersal syndrom animal species  7240652
291Germination efficiency  1238706595
295Seed oil content  452602882
297Seed protein content  360302308
299Seed storage behaviour  12492010291
304Fruit type  390303691
305Species diaspore type  13975483696
309Leaf inferior epiderm thickness  87022
310Leaf superior epidermis thickness  87022
311Leaf mesophyll thickness  87022
312Leaf mid vein thickness  1213450
313Leaf mid vein support tissue thickness  87022
314Stomatal conductance per leaf dry mass  122057
328Leaf volume of chloroplasts  94694
329Leaf number of chloroplasts per mesopyll cell  94694
330Leaf mesophyll area per leaf area  11517108
332Leaf mesophyll density  94694
334Net assimilation rate (NAR) 13790621
335Leaf area ratio (LAR)  42835219
336Leaf weight ratio (LWR)  1851435713
337Transpiration per leaf area  1473668
341Leaf boundary layer conductance  111032
344Germination lag time  63063
345Germination maximum rate per day  63063
346Leaf dry matter concentration  17017
347Leaf structural dry matter concentration  17017
348Root dry matter concentration  17017
349Whole plant dry matter concentration  17017
351Leaf structural dry matter content  17017
352Root dry matter content per fresh mass (RDMC)  66032
353Whole plant dry matter content  17017
354Leaf total volume  28028028
355Leaf intercellular volume  28028028
356Leaf tissue volume  28028028
357Leaf ratio of intercellular volume to leaf total volume  497336117
360Relative growth rate aboveground (RGR_above)  117084
368Specific leaf area (SLA) per fresh weight 28462754109
369Leaf soluble phenol content per dry mass  484848
371Whole plant aboveground vegetative dry mass 239239156
372Whole plant stem dry mass  12591259315
373Whole plant leaf dry mass  646646219
374Whole plant dry mass of reproductive structures 585585205
375Whole plant seed number  838381
376Whole plant seed mass  16416435
378Onset of seed maturation  303030
388Instantaneous water use effinciency  898988
389Instantaneous nitrogen use efficiency (PNUE)  433643
390Ratio of stem mass to whole plant mass  20135156
391Ratio of branch mass to whole plant mass  353535
392Ratio of wood mass to whole plant mass  440440114
395Canopy height observed 40683215777577
404Releasing height 202986292963
407Seed number per reproducton unit  1474401676
411Shoot growth form  716711333106
412Shoot branching  48796502990
414External support structure to stabilize the shoot  49341952845
444Leaf perimeter length  1082108279
445Leaf petiole length  828828150
446Leaf blade length  112377974839
447Leaf blade width  115237524679
448Plant height observed 25052505199
455Leaf carbon/nitrogen (C/N) ratio 26331593852
456Leaf phenolics content  471471229
457Leaf tannins content  471471229
458Mean distance between ramets  570570525
465Litter carbon/nitrogen (C/N) ratio  20234124
467Leaf carbon/phosphorus (C/P) ratio  414141
468Leaf palatability  271971979
470Bud source  3700168
471Leaf shape  311602980
472Age of maturity of resprouts  5750554
473Age of maturity of saplings  7870437
475Propagule  522202677
476Resprouting capacity after clipping  113075
477Resprouting capacity after disturbances  97061
478Resprouting capacity after fire  245143820
480Seed bank location  252501607
481Seed bank longevity  1109901509
482Post fire seedling emergence  21620746
483Post fire seedlings survival  70042
485Leaf fresh weight 28262798513
486Leaf chlorophyll content per dry mass  1251729
487Stem carbon/nitrogen (C/N) ratio 245245173
488Coarse root carbon/nitrogen (C/N) ratio 464646
489Fine root carbon/nitrogen (C/N) ratio 1054570
502Onset of flowering season  452717423034
503Reproductive plant height 820820271
504Vegetative plant height  962962308
512Leaf + paracotyledon area ratio (LPAR)  196097
513Leaf + paracotyledon weight ratio (LPWR)  196097
515Conduit (vessel and tracheid) density 14946517259
516Conduit (vessel and tracheid) area 3050512993
517Leaf area per sapwood area 515151
518Conduit lumen area per sapwood area 2280512271
519Phylogeny  448413483543
530Needle-leave form  26626042
531Leaf volume to area ratio  22522521
532Leaf total to projected area ratio  15515512
533Leaf mesophyll cell heigth  252511
534Leaf mean cell volume  50157
535Leaf volume fraction palisade to spongiophyll  371371223
536Leaf volume fraction of palisade parenchym  579418274
537Leaf volume fraction of spongiophyll  548387269
538Leaf volume fraction of epidermis  723556410
539Leaf volume fraction of hypodermis  164358
540Leaf epidermis cell size  2907674
541Leaf epidermal wall thickness  158158129
542Leaf nitrogen content per total area  24241
547Vmax per dry mass at ambient temperature 661462104
549Vmax per dry mass at 25 C 676726
550Vmax per leaf area at 25 C 757527
551Fraction of nitrogen in Rubisco 171713
552Leaf osmotic potential at full turgor  12112121
553Leaf osmotic potential at turgor loss  323212
554Leaf elastic modulus  424215
556Leaf apoplastic water concentration  331
559Leaf mass per area (LMA) of total leaf area  15314715
562Seed morphological adaptation to dispersal syndrom  2010184
563Disperal measurement distance  26600130
564Seed dispersal measurment relative abundance  26600130
565Species distribution and nativity status  2265408895
566BioFlor Phylogeny code  280802807
567Strategy type according to Grime  275102750
568Urbanity  280802807
569Hemeroby  926202831
570Number of floric zones  265702657
571Climbing mode  74074
573End of flowering season  2828432786
574Heterotrophic nutrition mode  788505754
576Metamorphoses for vegetative dispersal  142401423
577Apomixy  432503652
578Insemination autogamous or xenogamous  277702717
580Flower class  631703565
582Flower type  356303562
583Flower color  589002894
585Species reproduction  366803665
586Flower heterostyly  56043
588Flower pollen ovule ratio  3020238
589Self-incompatibility  309502630
590Flower sex timing  291102405
591Flower sex  418903631
592Flower self-incompatibility mechanism  5130502
593Flower UV reflecance in periphery  9270906
594Flower UV reflecance in centrum  9270906
595Flower UV reflectance pattern  9270906
597Species continentality  1146402782
598Species origin zonal  272601426
599Species origin altitude  272601426
600Chromosoms base number  382303083
601Chromosoms ploidy  382303083
602Chromosoms number  382303083
603Chromosoms cDNA content  11480684
604Chromosoms number  11480684
605Metamorphoses of shoot  280102407
607Metamorphoses for storage  32342962579
608Tolerance to mowing  6670667
609Tolerance to grazing  6670667
610Tolerance to trampling  6670667
612Sociological group  747702964
614Dispersal unit kind  000
615Dispersal unit heteromorphy  472501473
616Dispersal unit dry weight  36140882
617Seed heteromorphy  353001165
618Dispersal unit thickness  600001042
619Dispersal unit width  691801188
620Dispersal unit length  851101438
621Seed thickness  46050888
622Seed width  7764481597
623Dispersal unit appendix nutritional  107093
624Dispersal unit appendix laminar  202
625Dispersal unit appendix elongate  45036
626Dispersal unit additional cover  20020
627Seed appendix nutritional  1840184
628Seed appendix laminar  303
629Species mode of introduction  10170916
630Species neophyt in region since  3120270
631Species region of endemism and degree of naturalization in Germany  462403671
635nitrogen mineralisation rate  450450110
636Root exudation  36309
637Root porosity  25110759
638Alcohol Dehydrogenase (ADH) activity  14406
641Bark density  737737278
642Leaf Al content per dry mass 369115661180
643Leaf B content per dry mass  897897323
644Leaf Ba content per dry mass  2604897836
645Leaf Ca content per dry mass 376116121199
646Leaf Co content per dry mass  2404415741
647Leaf Cr content per dry mass  1614421550
648Leaf Cu content per dry mass  923904340
649Leaf Fe content per dry mass 313510961042
651Leaf Mg content per dry mass 349116121175
652Leaf Mn content per dry mass 327410921057
653Leaf Mo content per dry mass 571421221
654Leaf Na content per dry mass 318610961064
655Leaf Ni content per dry mass  944902356
656Leaf S content per dry mass 10981098405
657Leaf Se content per dry mass  666335289
658Leaf Si content per dry mass 10921092402
659Leaf Sr content per dry mass  2303447787
660Leaf Ti content per dry mass  903903325
661Leaf V content per dry mass  589416247
662Leaf Zn content per dry mass 329010971071
664Jmax per dry mass at ambient temperature 41123771
665Jmax per leaf area at ambient temperature 1073502219
666Light use efficiency as CO2 uptake per absorbed light  50445
667Light use efficiency as O2 release per absorbed light  23014
668Light use efficiency as CO2 uptake per incident light  47020
669Q10 of leaf respiration  239056
702Growth ring distinction  820805231
703"Wood porosity (ring-porous diffuse porous)"  705905221
704Vessel arrangement  13080923
706Vessel grouping  241501922
707Solitary vessel outline  251802178
708Vessel perforation plates  882805229
709Intervessel pits: arrangement and size  2211305176
710Vessel vestured pits  226001522
711Vessel - ray pitting  1039005225
712Vessel helical thickenings  18520712
713Mean tangential diameter of vessel lumina 978305224
715Vessel element length 1468805780
716Tyloses and deposits in vessels  276601925
717Wood vesselless  505
718Tracheids and fibres  7820692
719Ground tissue fibres  924205228
720Septate fibres and parenchyma-like fibre bands  753705229
721Fibre wall thickness  820105042
722Mean fibre lengths  1231905071
723Axial parenchyma  8730765
724Apotracheal axial parenchyma  349102192
725Paratracheal axial parenchyma  989103552
726Banded parenchyma  484502434
727Axial parenchyma cell type / strand length  1283504891
728Wood ray width  799705178
732Cellular composition of rays  1065705213
733Sheath cells  7710682
734Tile cells  66056
735Perforated ray cells  6710603
736Disjunctive ray parenchyma cell walls  5640513
737Rays per millimetre  751205172
738Wood rayless  20019
739Storied structure  18170727
740Oil and mucilage cells  4380213
741Intercellular canals  6440410
742Tubes  1790148
743Cambial variants  1700137
744Prismatic crystals  1028502717
745Wood druses  228088
746Other crystal types  9480421
747Other diagnostic crystal features  5580417
748Wood silica content  11560408
750Wood habit  738305229
751Wood of commercial importance  255701962
752Wood specific gravity  500402963
753Heartwood colour  911403027
754Wood odour  3110238
755Heartwood fluorescent  54030
756Water & ethanol extracts: fluorescence & colour  459099
757Froth test  34021
758Chrome Azurol-S test  302
759Burning splinter test  49031
776Regeneration after fire by seeds and/or resprouts  29102112781
780Tolerance to frequent fires  269902576
781Tolerance to infrequent fires  269702574
782Plant age at first flowering (primary juvenil period)  6620653
784Recommended minimimum fire interval  95094
785Seedbank longevity 3640353
786Flowering season  161723847253
790Fire regeneration category  830302638
801Dispersal distance  83083
806Vital attributes of persistence and establishment  454902389
807Evolutionary history (naturally / anecophytic)  280802807
808Leaflet length  29463321405
809Leaflet width  31933801409
816Maximum height observed of flower 2121766265
817Maximum height observed of leaf 2266875307
818Canopy diameter along the longest axis  1961596239
819Canopy diameter perpendicular to first axis  1929564232
820Leaf aspect ratio (width/length)  10221022174
821Leaf form coefficient (perimeter/area)  908908101
822Stem length  580580172
823Stem diameter  625625121
824Stem density  625625176
825Number of ramets  939392
826Clonality of ramets  230226199
827Canopy size  284284106
828Leaf hair length  909090
829Stem hair length  909090
830Leaf form index (area/length^2)  909090
831Coefficient of variation of plant height  909090
845VegClass PFT classification  20687206874333
846Family  26557219097403
856Leaf chlorotype  20659206594332
857Frost hardiness leaf / bud  713941
858Frost hardiness shoot  703
859Frost hardiness stem  1605
860Frost hardiness shoot cambium  201
861Frost hardiness twig  201
862Frost hardiness root  603
863Frost hardiness fine root  603
864Frost hardiness root cambium  201
865Frost hardiness seedling  201
869Ash content  20020
870Succulence  986791
872Lateral spread  99068
873Reproductive phenology  62031
874Seed number per ramet  28370586
875Occurrence of seed hard coats or seed transport by ants  31031
876Hairs orientation  505
877Mean epidermal cell area  54054
878Shoot (above ground) biomass nitrogen content per dry mass  64909
879Shoot (above ground) biomass phosphorus content per dry mass  51005
880Clonal growth form  2450132
881Aerenchyma  7160569
882Plant life form (Raunkiaer life form)  64936554767864
883Type of vegetative regeneration  4400321
884Regeneration capacity after damage  2020118
885Physical defense structures  1400120
886Chemical defense structures  8880366
887Seed shedding season  53692441788
888Mating system  179901304
889Seed height  12560388
890Seed shape  568567524
891Seed morphology  15870851
892Seed or dispersal unit floating  7280667
893Digestion  101
894Number of seeds per dispersal unit  150047
895Xerochasy/Hydrochasy  3220304
896Germination temperature  17400601
897Germination or establishment temperature sensiticity  1280104
898Dormancy  1257110820
899Season of seedling emergence / germination  22551361381
900Germination requirement  5280379
901Light requirement  7270469
902Clonal growth organ  4690274
903Role of clonal growth organ in plant growth  59032
904Shoot life span  35022
905Persistence of connection between clonal growth organs  46027
906Individual life span  23012
907Frequency of seeding  41034
908Bud frequency per bud bank layer  9390331
909Number of offspring shoots per parent shoot per year  45024
910Seed ripening season  192096
912Roots swollen  11811816
913Roots with nodules  11811816
914Stem volume  19511892
915Root dry mass  12312317
916Shoot dry mass  12312317
917Height of lowest branch  11711716
918Legume  626039
919Leaf lamina cell size palisade parenchyma  150051
920Leaf lamina cell size abaxial (lower) epidermis (hypodermis)  155053
921Leaf vascular bundle volume fraction: xylem  160055
922Leaf vascular bundle volume fraction: phloem  160055
923Leaf vascular bundle volume fraction: sclerenchyma  160055
924Leaf vascular bundle cell size: xylem  74033
925Leaf vascular bundle cell size: phloem  89037
926Leaf vascular bundle cell size: sclerenchyma  25011
927Leaf vessel maximum diameter  160055
928Leaf fraction sclerified tissue midrib  143051
929Leaf fraction sclerified tissue lamina  117043
930Stem vessel maximum diameter  108080
931Root vessel maximum diameter  13013
932Stem fraction cell wall in xylem (CWx)  47047
933Stem fraction cell wall in pith  404
934Stem fraction cell wall in collenchyma  606
935Stem area pith  80079
936Stem area xylem  80079
937Stem area other tissues but pith and xylem  80079
938Stem total area of cross section  80079
939Root area xylem  12012
940% pith  80079
941% xylem  80079
942% tissues other than pith or xylem  80079
943% collenchyma  19019
944% of total area - xylem cell wall  48048
945% of total area - collenchyma cell wall  32032
946% of total area - pith cell wall  32032
947Total % cell wall  32032
948Stem dry weight  78077
949Stem height (mm)  78077
952Root weight fraction (RWF or RMR or RWR) 1660121
953Leaf weight fraction (LWF or LMR) 1380105
954Paracotyledon weight fraction (PWF)  68068
956Paracotyledons dry weight  89088
957Phosphorus uptake rate  23023
959Multi-stemness  214214213
960Seasonality of growth  330330329
961Diaspore size (diameter)  484848
962Diaspore color  474747
963Number of seeds per dispersal unit  484848
964Internode length  828272
965Specific internode mass  828272
966Specific petiole length  818172
967Internode mass to metamer fraction  828272
968Petiole mass to metamer fraction  828272
969Leaf mass to metamer fraction  828272
970SLA of leaf lamina 787870
971Leaf area to metamer ratio  828272
972Leaf lamina density  787870
973Internode area to leaf area ratio  828272
974Internode density  828272
975Internode area  828272
983Whole pant aboveground biomass 563563275
984Whole pant aboveground necromass  12612688
985Leaf inclination  228228120
986Stem nitrogen content 675452170
987Stem carbon content 266198147
988Whole plant aboveground nitrogen pool vegetative  196196125
989Whole plant aboveground carbon/nitrogen (C/N) ratio (vegetative)  192192126
990Whole plant leaf area  220220127
992Branch wood density  727265
993Nitrate reductase activity  747466
994Leaf chlorophyll a content per area  777768
995Leaf chlorophyll b content per area  777768
996Leaf chlorophyll a+b content per area  43542785
997Leaf chlorophyll a/b content  818169
998Leaf Xanthophyll + Carotene.  000
999Leaf ratio of total Chlorophyll / Carotene  000
1000Electron Transport Rate  777768
1001Vessel scalariform perforations  10750939
1002Vessel bars per perforation plate  31520918
1003Vessel diameter 32090939
1004Fibre length  30230892
1005Leaf and fine root turnover  129026
1006Fine root labile compound content  12012
1007Fine root cellulose content  12012
1008Fine root lignin content  12012
1009Dead wood lignin content  30026
1010Dead wood cellulose content  30026
1011Leaf water potential at initial reduction of stomatal conductance  40034
1012Leaf water potential at final reduction of stomatal conductance  40034
1013Vapor pressure deficit at initial reduction of stomatal conductance  37034
1014Vapor pressure deficit at final reduction of stomatal conductance  37034
1015Litter labile compound content  102071
1016Dead wood carbon to nitrogen ratio  37032
1017Ratio of leaf area index of the whole leaves to projected leaf area index  1104
1018Light extinction coefficient  54028
1019New live wood carbon to new total wood carbon allocation  16016
1020New coarse root carbon to new stem carbon allocation  102023
1021New fine root carbon to new leaf carbon allocation  70041
1022New stem carbon to new leaf carbon allocation  247060
1023Water interception coefficient  504
1024Number of non-mature (growing) leaves per shoot  434315
1025Number of mature leaves per shoot  434315
1026"Vegetative plant height not elongated" 434315
1027"Flowering plant height heighest leaf elongated" 434315
1028"Flowering plant height heighest leaf not elongated" 434315
1029Height of inflorescence top  434315
1034Fraction of NPP allocated to branches  111
1035Fraction of NPP allocated to coarse root  111
1036Fraction of NPP allocated to fine roots  111
1037Fraction of NPP allocated to leaf  111
1039Intercept of respiration rate vs nitrogen content - mass basis - for different organs combined  12131
1040Slope of respiration rate vs nitrogen content - mass basis - for different organs combined  12121
1041Quantum yield of photosynthesis (Blackman function)  36363
1042"Light-saturated rate of photosynthesis at the growth CO2 concentration on a projected area basis"  26926516
1043Intercept of relationship between light-saturated photosynthetic rate (at the growth CO2 concentration) and leaf nitrogen content on a leaf area basis  10104
1044Slope of a relationship between light-saturated photosynthetic rate (at the growth CO2 concentration) and leaf nitrogen content on a leaf area basis.  10104
1045Rate of photosynthesis at saturating light and CO2 conditions  32322
1046Fraction of NPP allocated to stem  111
1047"Branch maintenance respiration at 15¡C on area basis from exponential regression"  12123
1048"Branch maintenance respiration at 15¡C on area basis from linear regression"  12123
1049"Branch total respiration at 15¡C on area basis from exponential regression"  13131
1050"Branch total respiration at 15¡C on area basis from linear regression"  13131
1051"Slope of linear regression of branch total respiration on area basis vs. Temperature"  13131
1052Mean date of budburst  661
1053Ratio of branch nitrogen concentration to foliage nitrogen concentration  111
1054Intercept of branch total respiration rate (growing tissue) vs branch nitrogen content - mass basis  221
1055Slope of branch total respiration rate (growing tissue) vs branch nitrogen content - mass basis  221
1056"Branch maintenance respiration at 15¡C on mass basis from exponential regression"  12123
1057"Branch maintenance respiration at 15¡C on mass basis from linear regression"  12123
1058"Branch total respiration at 15¡C on mass basis from exponential regression"  13131
1059"Branch total respiration at 15¡C on mass basis from linear regression"  13131
1060"Slope of linear regression of branch total respiration on mass basis vs. Temperature"  13131
1061"Branch total respiration at 15¡C on nitrogen basis from exponential regression"  13131
1062"Branch total respiration at 15¡C on nitrogen basis from linear regression"  13131
1063"Slope of linear regression of branch total respiration on nitrogen basis vs. Temperature"  13131
1064"Branch maintenance respiration surface area basis at the given temperature"  12123
1065"Branch total respiration (growing) surface area basis at the given temperature"  38384
1066"Branch maintenance respiration mass basis at the given temperature"  12123
1067"Branch total respiration (growing) mass basis at the given temperature"  42425
1068Branch wood carbon concentration  10101
1069Branch dry weight per unit ground area  26262
1070Branch dry weight per plant  1161169
1071Branch nitrogen concentration  27274
1072"Branch total respiration (growing) nitrogen basis at the given temperature"  24241
1073Branch total nitrogen content per unit ground area  661
1074Branch total nitrogen content per plant  331
1075"Branch total respiration (growing) volume basis at the given temperature"  221
1076Exponential coefficient of branch total respiration vs temperature  41412
1077Clumping factor: the ratio of shoot projected area to leaf projected area (for coniferous species)  663
1078Coarse root total nitrogen content per unit ground area  661
1079Coarse root carbon concentration  441
1080Coarse root dry weight per unit ground area  26263
1081Dry weight of coarse roots per plant  64645
1082Coarse root nitrogen concentration (dry weight basis)  14144
1083Diameter at base (seedlings)  1411416
1084Diameter at base of crown (seedlings)  73733
1085Constant K in nitrogen uptake equation (functional balance model)  221
1086Functional balance constant: ratio of nitrogen uptake to NPP  221
1087Root activity: rate of uptake of nitrogen per unit root biomass (in functional balance model)  221
1088Fraction of total plant DW in branch  552
1089Fraction of total plant DW in coarse roots  10104
1090Fraction of total plant DW in fine roots  10104
1091Fraction of total plant DW in foliage 25254
1092Total fine root nitrogen content per unit ground area 661
1093Total fine root nitrogen content per plant 221
1094Fraction of total plant DW in root (coarse + fine)  34344
1095Fine root carbon content  141141134
1096Dry weight of fine roots per unit ground area  30304
1097Dry weight of fine roots per plant  62625
1098Annual turnover of fine roots (fraction of total fine root biomass lost per annum)  221
1099Fine root nitrogen concentration (dry weight basis) 153145137
1100Fraction of stem wood volume which is live sapwood  882
1101Fraction of total plant DW in stem  552
1102Fraction of total plant DW in wood (stem + branch)  35355
1103CO2 compensation point  26264
1104Fraction of ground area covered by projected canopy  42424
1105Green crown length  424210
1106Stomatal conductance when irradiance tends to zero in the Ball-Berry model  32328
1107"Slope in the Ball-Berry model assuming that soil water content does not affect stomatal conductance directly."  38388
1108Constant in the linear response of stomatal conductance to CO2 (Jarvis model)  662
1109Constant in the response of stomatal conductance to VPD (Jarvis model)  16166
1110Value of soil water potential (negative) above which stomatal conductance is unaffected by soil water content (Jarvis model)  442
1111Value of soil water potential (negative) below which stomatal conductance is zero (Jarvis model)  442
1112Constant D0 in the Leuning model.  40404
1113Stomatal conductance when the irradiance tends to zero in the Leuning model  40404
1114"Slope in the Ball-Berry-Leuning model under the assumption that stomatal conductance is independent of water stress."  42424
1115"Stomatal conductance to water vapour under conditions of light saturation high relative humidity and growth CO2 concentration."  18418015
1116"Stomatal conductance in the limit of saturating light zero VPD maximum soil water content and at the reference value of Cs = 350 ppm"  26267
1117Average tree height 21621613
1118Proportion of rainfall intercepted by canopy  111
1119"Potential electron transport rate on a projected area basis"  16716315
1120Intercept of Jmax (potential rate of electron transport) versus leaf nitrogen content (on a leaf area basis)  28287
1121Slope of Jmax (potential rate of electron transport) vs. leaf nitrogen content (calculated on a leaf area basis)  28287
1122Activation energy of electron transport  10103
1123Parameter H in the temperature response of electron transport (J)  10103
1124Parameter S in the temperature response of the electron transport rate J  10103
1125Canopy radiation extinction coefficient  663
1126Proportion of leaf area in given age class (age class should be specified in category field)  12122
1127"Leaf area index projected of a single canopy layer. Indicate height of layer with Category field."  90903
1128"Leaf maintenance respiration at 15¡C on area basis from exponential regression"  881
1129"Leaf respiration at 15¡C on area basis from exponential regression"  11112
1130"Leaf respiration at 15¡C on area basis from linear regression"  771
1131"Slope of linear regression of leaf total respiration on area basis vs. Temperature"  771
1132"Leaf area projected per crown"  82827
1133"Leaf maintenance respiration area basis at the given temperature."  34343
1134"Leaf respiration (growing tissue) area basis at the given temperature"  69697
1135Leaf carotenoid content  441
1136Intercept of leaf chrorophyll content (projected area basis) vs. leaf nitrogen content (area basis).  331
1137Slope of leaf chlorphyll content (projected area basis) vs. leaf nitrogen content (area basis)  331
1138Dry weight of foliage per unit ground area  32323
1139Dry weight of foliage per plant  15815811
1140Growth respiration coefficient for foliage  221
1141Annual turnover of foliage (fraction of total leaf biomass lost per annum)  221
1142Total leaf mineral concentration (obtained by summing concentrations of all major minerals)  17172
1143Maintenance respiration coefficient for foliage  221
1144"Leaf maintenance respiration mass basis at the given temperature"  28282
1145"Leaf maintenance respiration nitrogen basis at the given temperature"  28282
1146"Leaf respiration (growing tissue) nitrogen basis at the given temperature"  26262
1147Total canopy nitrogen content per unit ground area  661
1148Total leaf nitrogen content per plant  442
1149Leaf protein content  69412
1150Leaf Rubisco content (projected area basis)  48481
1151Leaf starch content  30715629
1152Leaf sugar content  30710733
1153Leaf total non-structural carbon concentration (TNC) 2427136
1154Proportion of leaf mass in given age class (age class should be specified in category field)  15151
1155Intercept of the relationship between leaf mass per area (LMA) and leaf nitrogen content.  20204
1156Slope of the relationship between leaf mass per area (LMA) and nitrogen content  20204
1157Intercept of leaf total respiration rate (growing tissue) vs leaf nitrogen content - mass basis  111
1158Slope of leaf total respiration rate (growing tissue) vs leaf nitrogen content - mass basis  111
1159"Leaf maintenance respiration at 15¡C on mass basis from exponential regression"  881
1160"Leaf respiration at 15¡C on mass basis from exponential regression"  992
1161"Leaf respiration at 15¡C on mass basis from linear regression"  771
1162"Slope of linear regression of leaf total respiration on mass basis vs. Temperature"  771
1163"Leaf maintenance respiration at 15¡C on nitrogen basis from exponential regression"  881
1164"Leaf respiration at 15¡C on nitrogen basis from exponential regression"  771
1165"Leaf respiration at 15¡C on nitrogen basis from linear regression"  771
1166"Slope of linear regression of leaf total respiration on nitrogen basis vs. Temperature"  771
1167Fraction of leaf nitrogen which is retranslocated prior to leaf senescence  331
1168Exponential coefficient of leaf maintenance respiration vs temperature  881
1169Exponential coefficient of leaf total respiration vs temperature  25252
1171Intercept of the relationship betweeen leaf nitrogen and leaf mass per area (LMA)  19194
1172Slope of the relationship betweeen leaf nitrogen and leaf mass per area (LMA)  19194
1173Number of branches per tree  71716
1174"Leaf mitochondrial respiration in the light calculated by fitting the Blackman response"  16162
1175"Leaf mitochondrial respiration in the light calculated by fitting the Farquhar-von Caemmerer model to the A-Ci curve"  11711310
1176Ratio of root nitrogen concentration to foliage nitrogen concentration  332
1177"Root maintenance respiration at 15¡C on mass basis from exponential regression"  12123
1178"Root maintenance respiration at 15¡C on mass basis from linear regression"  12123
1179"Root respiration at 15¡C on mass basis from exponential regression"  12121
1180"Root respiration at 15¡C on mass basis from linear regression"  12121
1181"Slope of linear regression of root total respiration on mass basis vs. Temperature"  12121
1182"Root respiration at 15¡C on nitrogen basis from exponential regression"  12121
1183"Root respiration at 15¡C on nitrogen basis from linear regression"  12121
1184"Slope of linear regression of root total respiration on nitrogen basis vs. Temperature"  12121
1185"Root respiration (growing tissue) area basis at the given temperature"  221
1186Dry weight of total (fine + coarse) roots per unit ground area  331
1187Dry weight of total (fine + coarse) roots per plant  686810
1188"Root maintenance respiration mass basis at the given temperature"  18185
1189"Root respiration (growing tissue) mass basis at the given temperature"  42425
1190"Root respiration (growing tissue) nitrogen basis at the given temperature"  24242
1191Exponential coefficient of root total respiration vs temperature  24241
1192Woody area index  11115
1193"Stem maintenance respiration at 15¡C on area basis from exponential regression"  14141
1194"Stem respiration at 15¡C on area basis from exponential regression"  14141
1196Intercept of stem nitrogen concentration vs foliage nitrogen concentration  331
1197Slope of stem nitrogen concentration vs. leaf nitrogen concentration  111
1198"Stem maintenance respiration at 15¡C on mass basis from exponential regression"  16162
1199"Stem respiration at 15¡C on mass basis from exponential regression"  22222
1200"Stem maintenance respiration at 15¡C on nitrogen basis from exponential regression"  14141
1201"Stem respiration at 15¡C on nitrogen basis from exponential regression"  16162
1202Exponential coefficient of Stem maintenance respiration vs temperature  18182
1203Exponential coefficient of Stem total respiration vs temperature  24242
1204"Stem maintenance respiration surface area basis at the given temperature"  16161
1205"Stem respiration (growing) surface area basis at the given temperature"  16161
1206Stem wood carbon concentration  1264126437
1207Dry weight of stem per unit ground area  28282
1208Dry weight of stem per plant  1001008
1209Empirical form factor in calculating sapwood C  551
1210Growth respiration coefficient for stems  111
1211"Stem maintenance respiration mass basis at the given temperature"  14141
1212"Stem respiration (growing) mass basis at the given temperature"  14141
1213"Stem maintenance respiration nitrogen basis at the given temperature"  14141
1214"Stem respiration (growing) nitrogen basis at the given temperature"  14141
1215Total stem nitrogen content (per unit ground area)  661
1216Total stem nitrogen content per plant  331
1217Stem surface area to dry weight ratio  332
1218Stem surface area to volume ratio  111
1219"Stem maintenance respiration volume basis at the given temperature"  16161
1220"Stem respiration (growing) volume basis at the given temperature"  26261
1222Initial slope of total soil respiration response to soil water content  332
1223Q10 of total soil respiration  332
1224Asymptote of total soil respiration at high soil water content  332
1225Curvature of the light-response of RuBP regeneration  44445
1226Litter carbon concentration  222
1227Litter nitrogen concentration  222
1228Maximum temperature at which light-saturated photosynthesis (AMAX) is positive.  664
1229Minimum temperature at which light-saturated photosynthesis (AMAX) is positive.  664
1230Optimum temperature for light-saturated photosynthesis Amax  664
1231Average leaf transmittance of photosynthetically active radiation  111
1232"Maximum rate of Rubisco activity on a projected area basis"  16616215
1233Intercept of the relationship between maximum Rubisco activity and leaf nitrogen content (on a leaf area basis)  28287
1234Slope of the relationship between maximum Rubisco activity and leaf nitrogen content (on a leaf area basis)  28287
1235Activation energy of Rubisco activity  10103
1236Total above-ground woody biomass per plant  17174
1237Coefficient in allometric relationship for stem mass vs height & diameter  16165
1238Exponent in allometric relationship for stem mass vs height and diameter  16165
1239Leaf Cl content per dry mass  21010655
1240Leaf Sc content per dry mass  20900653
1242Leaf As content per dry mass  3346190
1243Leaf Br content per dry mass  21530666
1244Leaf Rb content per dry mass  20866651
1245Leaf Ag content per dry mass  126072
1246Leaf Cd content per dry mass  252674
1247Leaf Sb content per dry mass  12206381
1248Leaf Cs content per dry mass  15550527
1249Leaf La content per dry mass  17810558
1250Leaf Ce content per dry mass  14730491
1251Leaf Nd content per dry mass  3140124
1252Leaf Sm content per dry mass  18360576
1253Leaf Eu content per dry mass  10530404
1254Leaf Gd content per dry mass  249098
1255Leaf Tb content per dry mass  6120247
1256Leaf Dy content per dry mass  118057
1257Leaf Yb content per dry mass  6760271
1258Leaf Lu content per dry mass  4000156
1259Leaf Au content per dry mass  3450224
1260Leaf Hg content per dry mass  4936230
1261Leaf Th content per dry mass  9440383
1262Leaf U content per dry mass  50036
1263Leaf Hf content per dry mass  3070181
1264Leaf Ta content per dry mass  32027
1265Secretory elements in Wood  5840478
1266Silica [SiO2] bodies in wood  3010297
1267Furcation index  3313893313893860
1268Height of lowest branch versus total plant height  199199106
1269Root radial oxygen loss  43005
1271True leaf carbon isotope discrimination  442
1272Leaf carbon content per area  621949021864
1273Leaf physical strength: puncturability  227227159
1274Leaf physical strength: resistance to fracture  000
1275Vcmax per leaf area 1725977267
1276Leaf water content per area  323222
1277Leaf mesophyll cell size (area)  161055
1278Stomatal conductance per leaf dry mass  841019
1279Area at base  12212217
1280Federal T/E Status  61060
1281National Wetland Indicator  120201198
1282Active Growth Period  212702074
1283After Harvest Regrowth Rate  139801370
1284Bloat  194001897
1285Carbon/nitrogen (C/N) Ratio  192501877
1286Coppice Potential  218502130
1287Fall Conspicuous  218502130
1288Fire Resistant  218502130
1289Flower Color  210202053
1290Flower Conspicuous  218502130
1291Foliage Color  216702112
1292Foliage Porosity Summer  22261072102
1293Foliage Porosity Winter  210702053
1294Foliage Texture  24572962396
1295Fruit/Seed Color  207302022
1296Fruit/Seed Conspicuous  218502130
1297Growth Form  213202077
1298Growth Rate  211102057
1299Height at 20 Years  10250995
1300Known Allelopath  218502130
1301Leaf Retention  218502130
1302Lifespan  214002085
1303Low Growing Grass  218502130
1304Shape and Orientation  214002086
1305Toxicity  210602053
1306Adapted to Coarse Textured Soils  218502130
1307Adapted to Fine Textured Soils  218502130
1308Adapted to Medium Textured Soils  218502130
1309Anaerobic Tolerance  213902085
1310CaCO3 Tolerance  209202041
1311Cold Stratification Required  218502130
1312Fertility Requirement  214002088
1313Fire Tolerance  206002008
1314"Frost Free Days Minimum"  217502121
1315Hedge Tolerance  205502009
1316Moisture Use  203701988
1317"pH Minimum"  211002057
1318"pH Maximum"  211102057
1319"Planting Density per Acre Minimum"  166801627
1320"Planting Density per Acre Maximum"  166801627
1321"Precipitation Minimum"  211702062
1322"Precipitation Maximum"  211802063
1323"Root Depth Minimum"  216802115
1324Salinity Tolerance  211502062
1325"Temperature Minimum"  217402119
1326Commercial Availability  206402013
1327Fruit/Seed Abundance  200101952
1328Fruit/Seed Period Begin  208002029
1329Fruit/Seed Period End  207802028
1330Fruit/Seed Persistence  218502130
1331Propagated by Bare Root  218502130
1332Propagated by Bulb  218502130
1333Propagated by Container  218502130
1334Propagated by Corm  218502130
1335Propagated by Cuttings  218502130
1336Propagated by Seed  218502130
1337Propagated by Sod  218502130
1338Propagated by Sprigs  218502130
1339Propagated by Tubers  218502130
1340Seed Spread Rate  206302010
1341Seedling Vigor  198901938
1342Small Grain  218502130
1343Vegetative Spread Rate  203401983
1344Berry/Nut/Seed Product  218502130
1345Christmas Tree Product  218502130
1346Fodder Product  218502130
1347Fuelwood Product  218502130
1348Lumber Product  218502130
1349Naval Store Product  218502130
1350Nursery Stock Product  218502130
1351Palatable Browse Animal  166901619
1352Palatable Graze Animal  157301525
1353Palatable Human  218502130
1354Post Product  218502130
1355Protein Potential  125901229
1356Pulpwood Product  218502130
1357Veneer Product  218502130
1359Specific Root Length (m/g) of Fine Roots  245245191
1360Mean Flr Date (Julian day)  139139139
1361Flowering Period (Flr) Length (days)  139139139
1365Leaf pH  281281104
1367Wood Nitrogen content  1266126637
1368Wood phosphorus content  1056105635
1369Bark Carbon Content  43943927
1370Bark Nitrogen Content  43843827
1371Bark phosphorus Content  42542526
1372Coarse root phosphorus content per dry mass  111
1373Architectural allometries: stem diameter versus aboveground biomass  333
1374Fv/Fm  48482
1375Stem/Wood Ca content per dry mass  663
1376Stem/Wood Mg content per dry mass  663
1377Stem/Wood Al content per dry mass  663
1378Stem/Wood Fe content per dry mass  663
1379Stem/Wood Na content per dry mass  663
1380Stem/Wood K content per dry mass  663
1381Stem/Wood S content per dry mass  663
1382Stem/Wood Hg content per dry mass  663
1383Stem/Wood Sb content per dry mass  663
1384Stem/Wood Mo content per dry mass  663
1385Stem/Wood Cd content per dry mass  663
1386Stem/Wood Pb content per dry mass  663
1387Stem/Wood Ni content per dry mass  663
1388Stem/Wood Cu content per dry mass  663
1389Stem/Wood Ti content per dry mass  663
1390Stem/Wood Zn content per dry mass  663
1391Stem/Wood As content per dry mass  663
1392Stem/Wood Cr content per dry mass  663
1393Coarse Root Ca content per dry mass  111
1394Coarse Root Mg content per dry mass  111
1395Coarse Root Al content per dry mass  111
1396Coarse Root Fe content per dry mass  111
1397Coarse Root Na content per dry mass  111
1398Coarse Root K content per dry mass  111
1399Coarse Root S content per dry mass  111
1400Coarse Root Hg content per dry mass  111
1401Coarse Root Sb content per dry mass  111
1402Coarse Root Mo content per dry mass  111
1403Coarse Root Cd content per dry mass  111
1404Coarse Root Pb content per dry mass  111
1405Coarse Root Ni content per dry mass  111
1406Coarse Root Cu content per dry mass  111
1407Coarse Root Ti content per dry mass  111
1408Coarse Root Zn content per dry mass  111
1409Coarse Root As content per dry mass  111
1410Coarse Root Cr content per dry mass  111
1425Gymnosperm/Angiosperm/Tree/Herb  2130267279
1426Leaf respiration per dry mass at 20 C  1386267263
1427Root respiration at measurement temperature  744083
1428Root respiration calculated to 25C  744083
1429Root respiration calculated to 20C  744083
1430Root respiration calculated to mean temp of growth season  000
1433SLA per leaf fresh weight  172144159
1440Stem respiration rate  38023316
1446Leaf organic nitrogen content 145019
1447Leaf nitrate (NO3-) content 142019
1448Root sugar content per dry mass  48016
1449Root Respiration at 18C  15015
1450Root Respiration at 23C  46016
1451Root Respiration at 28C  16016
1452Leaf respiration per dry mass (Rdmass) at 18 C  15015
1453Leaf respiration per dry mass (Rdmass) at 23 C  45016
1454Leaf respiration per dry mass (Rdmass) at 28 C  16016
1455Leaf respiration per dry area (Rdarea) at 18 C  15015
1456Leaf respiration per dry area (Rdarea) at 23 C  45016
1457Leaf respiration per dry area (Rdarea) at 28 C  16016
1458Leaf photosynthesis at saturating light (Asat) mass 18 C  15015
1459Leaf photosynthesis at saturating light (Asat) mass 23 C  15015
1460Leaf photosynthesis at saturating light (Asat) mass 28 C  16016
1461Photosynthesis per leaf area at irradiance of 500 micro mol m-2 s-1  000
1462Leaf respiration in light per area  16010
1463Photosynthesis per leaf mass at irradiance of 500 micro mol m-2 s-1  16010
1464Leaf respiration in light per mass  32010
1465Ratio leaf respiration in light / respiration in the dark  16010
1466"Difference of leaf respiration in dark - respiration in the light mass based"  16010
1467Gamma star (Farquhar model)  16010
1468Farquhar model: Oxygenation capacity  16010
1469Cotyledon Nitrogen content  20010
1470Total nitrogen content (%) 20010
1471Root nitrogen ratio  20010
1472Stem nitrogen ratio  20010
1473Leaf nitrogen ratio  20010
1474Growth per nitrogen  20010
1475Growth per nitrogen in foliage  20010
1476Nitrogen uptake ratio  20010
1477Root carbon content (%)  000
1478Cotyledon Carbon content  20010
1479Root TNC  20010
1480Stem TNC  20010